on edge

In the film, the humans have amassed a giant armory of weapons, which makes the apes very on-edge, and leads to them revolting.

The front may be formed of two course of brick-on-edge, and if divisions are required, they should be formed in the same way.

If you want a real, first-class, tooth-on-edge Doone valley, the place to look for it is in the book.