Four columell curved, the two lateral ones prolonged downwards into two vertical, tuberculate feet of half their length.
Spores large (40–50 broad), globose or oblong, tuberculate.
The skin of the dorsum is thick and glandular, but not tuberculate.
Specimens from the Peace River, collected on August 10, 1952, include females that were mostly spent and tuberculate males.
Tuberculate males were collected in the Powder River on June 30, 1957.
The carnassial teeth are not typically developed, and the molars are broad and tuberculate.
Shell subspherical, tuberculate, with slight sagittal stricture and circular internal ring.
Shell nut-shaped, tuberculate, with subregular circular pores.
Shell nut-shaped, tuberculate, with sharp sagittal stricture.
Shell nut-shaped, tuberculate, without external sagittal stricture.