They have the stems of pennae, and the downy barbs and barbules of plumulae.
Compare as well Met V 294-98 'Musa loquebatur: pennae sonuere per auras, / uoxque salutantum ramis ueniebat ab altis.
Before them was spread a tablet covered with quills (pennae) which flew in the precepts of God.
(a) pennae including quills and coverts, (b) down feathers or plumulae, and (c) filoplumes which are rudimentary feathers.
In some birds, however, such as the Ratitae and the Penguins, pennae are evenly distributed over the whole body.
The pens used for writing were either made of reeds (calami) or of quills (pennae).